Andrena: select species diagnoses

Washinhgton, 10mm;
Clypeus dull, impunctate;
Tergae impunctate;
Complete corbicula;
Complete hair bands;
Scutum hair long;
Cuneate tibia;
Vertex ~1.4ocd;
Fovea dark/light;
Weak humeral angle

Andrena angustitarsata
Andrena frigida

LaBerge: "Andrena angustitarsata Viereck is an extremely common western Andrena, distinguishable in either sex by the terga being punctureless and opaque due to the dense regular tessellation."

Above: Clypeus sculpturing uniformly dull, impunctate.
Below: Humeral angle weak, pronotal ridge absent.

Above: Vertex ~1.4 ocd; fovea dark above.
Below: Clypeal hairs plumose.

Andrena astragali (Ptilandrena)

Shortcut: Andrena astragali forages exclusively on death camas, which is visited by no other bee.

Andrena auricoma (Ptilandrena)

The overall orange coloration is notable, as is the resemblance to A. astragali, another Ptilandrena.

Oregon, 9mm;
Clypeus dull;
labral process emarginate;

Triangle 0% sculptured;

Incomplete corbicula;
Complete hair bands;
T2 pitting 1-3;
Cuneate tibia;
Pale scopal hairs;

Andrena auricoma
Andrena bucculenta
Andrena lutehirta

DL has no images of bucculenta or lutehirta, so overall appearance is unhelpful. A. lutehirta is scored as having a long vertex (1.5-2od), whereas this specimen has vertex less than one ocellar diameter. This specimen has 3 SMC, whereas A. bucculenta is scored as having 2SMC, a condition that excludes the subgenus of the other two options. This leaves a diagnosis of A. auricoma.

Above: An 88 year old specimen from OSU. Note cuneate tibia.
Below: The sculpturing here is 0% sculptured, although the surface texture is borderline. The label here indicates Euandrena. But Ptilandrena is the updated classification.

Cautionary notes: Scoring for western Andrena species is often incomplete.

The LaBerge subgenus key would seem to indicate Ptilandrena as having highly plumose tibial scopae, whereas this specimen has simple hairs. However LaBerge himself contradicts this: I have images of a specimen, with det. by LaBerge, of another Ptilandrena (A. nigrihirta) with tibial scopa very similar to this specimen, with simple hairs.

Above: Slightly emarginate labral process.
Below: Note super-shiny tergae, sculpturing of frons.

Andrena caerulea (Ptilandrena)

Washington, 9mm;
Metallic integument throughout;
Tibial scopa highly plumose;
Hair bands absent;
Light hairs - tergae, head, episternum;
3 submarginal cells.

... yields A. caerulea in DiscoverLife. LaBerge: "Both sexes of A. caerulea can be separated from related bees by the metallic coloration, the short clypeus which is only obscurely punctate, and the highly plumose hairs of the tibial scopae, the propodeal corbicula and the dorsal femoral flocculus." The small size, light hairs and plumosity diffentiate this species from A. nigrocaerulea.

Above: Highly plumose tibial scopa, dark(ish) hairs.
Below: Trochanteral flocculus (and adjacent femur) with plumose hairs.

Andrena carlini (Melandrena)

Easy calls, based on 2 images at right:

Michigan, 13mm;
Clypeus shiny/dull;
Median impunctate band wide (vs vicina);

Scopal and cheek hairs dark;

Complete corbicula;
Scutm hair long; thorax pale above, dark below;
Metasomal hairs dark; hair bands absent.

Andrena carlini
Andrena milwaukeensis
Andrena thaspii
Andrena vicina

Fovea dark above (at least). Match for A. carlini, not for A. vicina (all pale). [But see Washington A. vicina below]

Above: Vertex=2ocd. Excludes milwaukeensis and thaspii.
Below: Very short malar space. A. vicina would be short or moderate.

An additional specimen at right.

The mining bees of Minnesota (unpublished manuscript by Joel Gardner): "Andrena carlini is one of the most common Melandrena in Minnesota and easily recognized in the female by the hair of the head and thorax being bright yellow-white dorsally and black ventrally. Most other Minnesota Melandrena have the head and thorax with completely light hair." The exception would be A. vicina. I can't rule that out for this specimen. LaBerge: differs from A. vicina by "a [more] distinct median clypeal impunctate line with crowded punctures on either side." The A. carlini flocculus, not visible in photos, would be complete, but partial for vicina.

Male Andrena carolina (Andrena s. str.)

Mandibles distinctly decussate, with weak tooth;

Michigan, 8mm;
Clypeus hairs sparse; unpitted midline, pits spaced 1-3

Labral process emarginate;
Cheek 2X eye width;
Metasomal hair bands absent;
F1 longer than F2, but not F2+3.

Andrena carolina
Andrena chippewaensis
Andrena fulgida

These 3 species cannot be distinguished in DL. A. carolina is far more abundant, and associated with blueberries, likely the case for this specimen.

Quadrangle longer than broad. Tawny hairs. Clypeus shiny-dull, punctate except midline. Decussate mandibles. Max palp exceeds galea by 2 segments.

Cheek 2X as wide as eye. From DL: With distinct angle of 90 degrees or more along lower rear margin breaking the curve from top to bottom.

Above, from DL: UNCOMMON: With longitudinal rugulae in basal half, tessellate or smooth at apex
Below, from DL: DL: weak tooth not projecting below level of mandible articulation.

Andrena chlorogaster (Micrandrena)

T2 pits absent, granular surface;

Oregon, 7mm;
Clypeus dull;
Integument metallic;

Pale hair bands, hairs pale,
Labral process entire;
Propodeal sculpturing 50+%;
Metasomal hair bands absent;
Corbicula incomplete.

Result: Andrena chlorogaster. A second approach, given subgenus Micrandrena: consult key by Ribble. A. chlorogaster is consistent.

Andrena crataegi (Plastandrena)

Oregon, 12mm;
Clypeus shiny/dull;

Triangle 100%;

Incomplete corbicula;
T2 pitting dense;
Simple scopal hairs;
Scutum shiny.

Andrena amphibola
Andrena crataegi

A. amphiloba does not match. We would expect dark hairs on the tibia, a normal tibial spur, and a fovea like Trachandrena.

Above: "Noted for coarse propodeum and curved and thickened inner hind tibial spur."
Below: Distinctive carinate ridge of hind femur--a crataegi thing.

Andrena crataegi (Plastandrena)

Shortcut: At a glance, this Michigan specimen is a look-alike to the preceding Oregon A. crataegi. Not shown here ..,. but it exhibits the same deep sculpturing of the delineated propodeal triangle-- a possible Trachandrena. One feature separates this one from any similars in Michigan: strongly curved hind tibial spur.

If I am right about both the OR and MI specimens, I can infer common features as typical of the species. E.g., remarkably orange tibial hairs.

Andrena cf. cressonii

See wilkella diagnosis. Many of the same traits apply. But evaluate wilkella vs cressonii:

T2 surface: granular for wilkella, shiny for cressonii;
Compare labral process images. Short and broadly truncate in this specimen.
Clypeus not flattened with this specimen,

Either of these conditions will narrow the result to A. cressonii, not wilkella. In fact, the summary description by Joel Gardner (on DL page) says this for A. cressonii: "is very easily recognized by ... the extremely short and wide labral process, and the shiny tergum with distinct, crowded punctures."

Another way that I get to A. cressonii: working through the subgenus key (In LaBerge, and Joel Gardener's key for MN mining bees), this specimen is a Holandrena. A. cressonii is the only member of the subgenus present in MI.

Shining tergae, incomplete corbicula, simple tibial scopa.

Clypeus dull, with unpitted median. Clypeus normally rounded.

A. cressonii-like labral process. Rounder clypeal punctures, unlike for wilkella.

Follow-up ... I re-examined to evaluate a couple of traits that required dismantling the specimen. These support the diagnosis of A. cressonii.

A. cressonii should have a "trochanteral flocculus incomplete, basal half with short hairs." The mess of hairs and pollen makes this a little hard to evaluate, but I think I see this in the photo.

Distinct humeral angle and pronotal ridge, in anterior view.

Distinct humeral angle and pronotal ridge, in lateral view.

Shiny surface, dense pitting of T2.

Andrena distans (Ptilandrena)

Michigan, 8mm, May;
Clypeus dull;
Labral process entire, long;
Tibia shape narrow;

Tibial hairs highly plumose, pale;
T2 hair bands nearly complete;
Scutum pitting 1-3;

A. distans
A. persimulata

There are several characters that will exclude A. persimulata. My specimen, A. distans, has no propodeal sculpturing, and "variable" trochanter hairs. Also: persimulata "Easily identified by the presence of a large projecting knob or large swelling at the bottom of the lateral face of the mesepisturnum."

Notable: Resemblance to A. erigeniae. Small, incomplete corbicula, plumose scopa.

The extra evidence: specimen collected from Geranium, on which A. distans is a specialist.

Clypeus dull, hairy, plumose. F1 long.
Labral process entire, long. Malar space short.

Above: DL scores T2 as granular, or shiny, and pits absent or present. Reasonable because this one is hard to call. The propdeal triangle - appears to me to have no, or weak and uniform sculpturing. Frizzy plumose hairs on narrow tibia.
Below: Vertex about 1 ocd. Scutum pits moderate. There is a humeral angle.

Andrena erigeniae (Ptilandrena)

Michigan, 9mm, April;
Clypeus dull;
Fovea narrow;

Tibial hairs highly plumose, pale;
T2 hair band absent;
T2 granular surface;
Propodeal triangle 5-50% scupltured

A. erigeniae

Notable: Slender, sparse hairs.

The key character is the unusual frizzy-plumose tibial scopa. Another determinative datum: the was specimen collected on spring beauty (Claytonia virginica) for which A. erigeniae is a specialist pollinator. Additional note: spring beauty has many other pollinators in the sites I follow.

Clypeus dull. Fovea narrow. NOTE: the fovea appear dark. This is partly a trick of the lighting. But also, LaBerge says: "facial foveae with tomentum often pale brown or yellow in upper half or more." The DL guide scores for pale, and "dark above, pale below."" I might add "all dark" to the scoring.

Above: DL scores T2 as granular, or shiny, and pits absent or present. Reasonable because this one is hard to call.
Below: frizzy-plumose hind tibia. On the flying insects, the reflectivity is noticeable.

Andrena erythronii (Leucandrena)

Michigan, 11mm, April;
Clypeus shiny/dull;
Weakly emarginate labral process;
F1>F2+F3;

Complete corbicula;

Tibial hairs simple, pale, long;
T2 granular surface;
Propodeal triangle ~20% scupltured

A. barbara
A. erythronii
A. thaspii

Notable: Abundantly hairy. Integument dark--more black than brown. Complete corbicula. Scopa pale + dark hairs. 11 mm length.

A priori, I believe this is A. erythronii. One, because it was collected in the vicinity of trout lily, a typical associate. Two, because this was suggested on iNat by Max McCarthy. Three, because a single unusual trait--flattened clypeus--is seen in MI only for A. erythronii and A. accepta (easily rejected).

In any case, I get A. erythronii and just two alternatives with the basic traits listed at left. Rule out A. barbara: "Pits on basal area of T2 distinct and crowded, separated by FAR less than 1 pit width ..." Rule out A. thaspii: "Labral process BIDENTATE."

So far so good: erythronii. However, DL can easily take me down a different path, if I choose trait conditions that I see in these images, that DL does not assign for A. erythronii. One is the faint but definite pitting on T2. However DL (and LaBerge) indicate an impunctate T2. Two, I see a nearly complete hair band on T2; DL scores erythronii as having broadly interrupted, or no T2 band, LaBerge says: "terga 2-4 with weak apical pale fasciae interrupted medially on terga 2 and 3."

If I have A. erythronii here, DL would probably fail to get a diagnosis. If this is NOT erythronii, DL can't say what it actually is.

Clypeus shiny, with wide impunctate band. Key: the central disk is flattened (confirmed with side view). True for only two MI species (if I choose "only" in the HAS menu option)

Long weakly emarginate labral process, with transverse striations.

F1>F2+F2.

Propodeal triangle, weakly sculptured.

T2 surface dull. Punctate? Hair band? See discussion.

Tibial hairs are lush, but simple.

Andrena evoluta (Diandrena)

Bidentate labral process;

Washington, 11mm;
Two submarginal cells;
Integument metallic;

Pale hairs

Result: Subgenus Diandrena, Andrena evoluta or A. chalybioides.

These two species are both reported as specialists on Agoseris sp (though this specimen was collected from Microseris laciniata, a different aster). According to LaBerge: "This species is most closely related to A. chalybioides from which it is distinguished by its more coarsely sculptured propodeal enclosure, darker colored anal fimbria, and lower and less sharply defined dorsolateral angles of the pronotum. These two taxa are allopatric and may later be found to be distinct only at the subspecies level." I don't have both species to compare, so I defer to H. Ikerd's determination.

Andrena forbesii (Trachandrena)

Triangle 100%, with enclosing carina;

Michigan, 10-12mm;
Clypeus shiny, impunctate midline;

Propodeum outside triangle roughly irregular;
Incomplete corbicula;
Hair bands absent;
Pale scopal hairs;
Scutum pitting sparser than in A. salicifloris;
Scutum texture dull;

Andrena sigmundi
Andrena miranda
Andrena rugosa

All matches are subgenus Trachandrena, as described for A. salicifloris. The list does not include A. forbesii, so I take issue with DL. My support for the A. forbesii diagnosis: 100 specimens from the Rufus Isaacs lab, many with J. Ascher dets.

Below: DL scores A. forbesii with ocellus-fovea distance greater than 1 od. In this pic, and close-up at right, the distance appears less than 1 od. If I ignore the DL score for fovea-ocellus distance, A. forbesii matches in DL.

Above: Clypeal impunctate medial line indistinct. Crowded punctures below ocelli, and weak rugulae. Also, normal hind tibial spurs--excludes A. sigmundi.

Above: Not miranda, described as unique among this group with "Propodeum with triangle without ridged sides, boundaries indistinct, adjacent areas often sculptured nearly as coarsely as triangle." My specimens: "propodeum distinctly finer outside than inside triangle" A. miranda or A. rugosa would also have tergum 2 with depressed apical area occupying three-fourths or more of midline" (about half for my specimens).

DL comments specific to A. forbesii: Face below ocelli duller than parocular area, with weak rugulae longitudinal or lacking, with punctures separated mostly by 0.5 puncture widths or less - Vertex above lateral ocelli equals 1.5 ocellar diameters or slightly less - Scutum hairs normal, long, erect, usually slender

Andrena cf. heraclei (trachandrena)

Strongly sculptured propodeum, scutum pits, coarse episternum = trachandrena; 12 species in MI.
The conundrum: The LaBerge Trachandrena key yields 2 choices in the second couplet, based on: "facial fovea extremely narrow below and at level of greatest separation from eye space between compound eye and fovea at least twice as broad as fovea at that level." The third couplet differentiates A. rugosa with "face above antennal fossa with distinct longitudinal rugae" from A. heraclei with "face above antennal fossa without rugae." Is this A. heraclei? DL offers A. ceanothi as having "Very end of facial fovea extremely narrow almost just a slot," which would seem to ALSO fit LaBerge couplet 2. In any case, I can get to A. heraclei in DL, with the following discrepancies: this specimen has T2 depressed area ~1/2 of tergum vs 2/3 - 3/4 for heraclei; AND this specimen with vertex less than 1 ocd vs 1.5 ocd for heraclei.

Note the narrow, parallel-sided hind tibia, with plumose scopa.

Very broad impunctate band on clypeus. I have other specimens where this is less pronounced.

Andrena imitatrix (Scrapteropsis)

Easy calls, based on 1st 3 images at right:

Michigan, 10mm;
Clypeus shiny/dull;

Triangle 100%;

Scutum w/ dense pitting;
T2 bands broadly interrupted;
T2 shiny, pit spacing 1-3.

Andrena alleghaniensis
Andrena imitatrix
Andrena morrisonella

Note on image: not clear here that clypeus is shiny in center. But it is.

Fovea broader than antennal socket: excludes A. alleghaniensis. Also: alleghaniensis with short scutal hairs.

Above: Labral process emarginate [alleghaniensis bidentate, morrisonella entire or weak].
Below: "Propodeal corbicula hairs in anterior half mostly or entirely branched [simple in morrisonella].

Andrena (Melandrena)

Michigan, 12mm, May;
Clypeus shiny/dull;
Vertex tall;

Tibial hairs dark;
T2 hair band absent;
T2 puncate, 1-3;
Propodeal triangle 50% scupltured

Abundance of caution. Consider also A. vicina, given that this specimen strongly resembles the diagnosis for that species.

A. nigrihirta
A. nivalis
A. regularis

Notable: Incomplete corbicula, tibial hairs long and dark, moderate malar space. The femur displays a slight blush of light hairs, precisely like the previous specimen.

My approach here is to start with: is this specimen NOT A. vicina? A. nivalis problems: scopal hairs (in the east) would be light; vertex would be 2 ocd height (this specimen a little less). A. nigrihirta would have pale/dark thoracic hairs (this specimen all pale). A. regularis would have hind femur hairs pale (this one mainly dark) and "Pygidial plate VERY NARROW forming and acute arrow-head shape" (this one with broad, truncate plate).

There is no certainty here. Another approach: use the Bouseman/LeBerge key for melandrena. This gets me, probably, to A. nivalis. The conservative name for this specimen: Andrena cf. vicina.

See A. vicina specimen. The head exhibits the same trait set. Narrower impunctate clypeal band. Not in photos: labral process entire, labrum with median crista.

See A. vicina specimen. The abdomen exhibits the same trait set.

Andrena miserabilis (Larandrena)

Easy calls, based on 1st 3 images at right:

Michigan, 8mm, May;
Clypeus shiny/dull;
Clypeus with unpitted median band;
vertex <1 ocd;

Labral process long, entire;

Hind tibia narrow;

The striking and unique (?) feature: clypeus with very wide unpitted band.

A. miserabilis

Note the narrow, parallel-sided hind tibia, with plumose scopa.

Very broad impunctate band on clypeus. I have other specimens where this is less pronounced.

White fovea, less than 1 ocd from ocellus. Short vertex.

Labral process entire, wide and long.

Andrena nigrihirta (Ptilandrena)

I can't examine this specimen, but have images I made of one at the Oregon State Arthropod Collection. The determination is by LaBerge/Ribble. Traits evident in the photos do NOT yield A. nigrihirta in Discoverlife ... unless I can be persuaded that lower episternal hairs are dark.

OSAC specimen above, USGS specimen below. Evidently the same bee, both with light thoracic hairs.

OSAC specimen above, USGS specimen below.

Andrena nigrocaerulea (Ptilandrena)

Oregon, 11mm;
Metallic integument throughout;
Clypeus dull;
Hair bands absent;
Dark hairs - tergae, head, episternum;
3 submarginal cells.

... yields A. nigrocaerulea in DiscoverLife. LaBerge: "The female of A. nigrocaerulea is readily distinguished from related species by the extensive dark body hairs, the complete lack of apical metasomal fasciae, the usually punctate and often shiny clypeus, and the metallic blue-black color of the integument." I assume here that "often shiny clypeus" does not preclude my specimens with a dull clypeus (which otherwise matches the image of the OSAC specimen below).

Images at right: A. nigrocaerulea specimen from Oregon State Arthropod Collection, determined by LaBerge/Ribble.

Reference: LeBerge specimen

I happen to have images of a western Oregon State University specimen of A. nivalis, with a Bouseman/LeBerge determination. This would likely be part of the material for their Melandrena key. Are these consistent with my unknown Melandrena above??

Andrena orthocarpi (Simandrena)

Clypeus dull, no median band;

Oregon, 7mm, June;
Fovea dark/light, broad;

Vertex 1 ocd;

Scutum pit spacing 1-3;
T2 hair band broadly interrupted;
T2 surface granular, pitting indistinct;

Tibial scopa pale, plumose;

Complete corbicula;
vein r shorter than 4 vein widths.

Result:

Below: Fovea long, malar space short, clypeal hairs plumose.

Andrena pallidofovea (Simandrena)

labral process entire and long;

Nevada, 9mm, June;
Clypeus dull;

vertex longer than 1-1.5 od;

Fovea more than 1 od from ocellus;

Complete corbicula;
Hind tibia cuneate, hairs pale and weakly branched;
T2 granular;

T2 hair band narrowly interrupted;

Result: Andrena pallidofovea

Below: Fovea long, malar space short, clypeal hairs plumose.

Andrena pensilis (Simandrena)

Fovea dark above, light below;

Oregon, 10mm;
Clypeus dull, without median band, flat in profile;
Maxillary palpus exceeds galea by two palpal segments;

Malar space extremely short;
Labral process entire;
Vertex ~1 ocd;
Pygidial plate w/ raised triangle;

Result: Andrena pensilis.

LaBerege: Andrena pensilis is a California species that is readily recognized in the female sex by the long, relative simple, tapering clypeal hairs, which are wavy distally (photo far right). This feature is evident in the half a dozen Oregon specimens I have.

Andrena NOT pensilis

Result: Andrena thaspii, A. pensilis, or A. lillooetensis.

Washington, May, 10mm;
Clypeus dull, without median band;
Labral process emarginate, long;
Vertex ~1ocd;
Malar space moderate;
scopal hairs long, simple, mixed;
T2 surface granular, pitting weak;
Hair bands broadly interrupted;
Incomplete corbicula;
Triangle sculpture less than 50%;
Tibia elongate, narrow;

A. lillooetensis appears as an artifact of limited scoring in DiscoverLife--it matches because potentially disqualifying traits are not recorded in the guide. Disqualifying for A. thaspii: "labral process bidentate with a basal transverse depression" and "malar space approximately ¼ width of mandible."

Disqualifying for A. pensilis: absense of wavy clypeal hairs. My specimen has a mostly hairless clypeus ... unlike 6 pensilis specimens that have pronounced wavy hairs (and entire labral processes). Also: the hind tibia shape of pensilis is close to cuneate, whereas this specimen has a narrow tibia. I confess that back-and-forth with Discoverlife gave me other "maybe, but ..." options. At one point I favored A. robertsonii. I am stumped.

Not pensilis: straight clypeal hairs, narrow hind tibia.

Not pensilis: emarginate labral process.

Andrena placata (Callandrena)

Michigan, 10mm, August;
Clypeus dull;
Labral process bidentate;

Tibial hairs simple;
T2 bands complete; pitting 1-3;
Fovea dark abve;
Propodeum 50-80%

A. gardineri
A. placata
A. wilkella

Evidence against A. wilkella: this specimen with plumose tibial scopa (simple in wilkella); absent impuncate center of clypeus (present in wilkella).
A. placata feature: "Terga 5 and 6 hairs usually dark brown."

Above: Medial impunctate band of clypeus absent; A. wilkella would have this condition.
Below: Distinctly bidentate labral process.

Above:Highly plumose tibial scopa; simple in wilkella.
Below:Of the three options, only A. placata has the middle leg basitatsus "expanded medially, broader than hind basitarsus.""

Andrena pruni (Melandrena)

Scopal and cheek hairs light;

Michigan, 12mm;
Clypeus shiny/dull;
Vertex long;
Median impunctate band wide;
Fovea light, narrow-ish;
Labral process entire, long;

Complete corbicula;
Scutm hair long; thorax pale hairs;
Scutum punctation dense;
Metasomal hairs light; hair bands absent;
Pygidial plate modified (excludes vicina and barbara).

Andrena pruni
Andrena dunningi

A. dunningi traits that don't match: Moderate or broad fovea, "UNIQUE in this group for having tibial scopal hairs below basitibial plate washed with black or dark brown."

Issues: interpretation of the fovea. A. pruni is distinguished from dunningi (and others) as UNIQUE in this group for having facial fovea separated from inner eye margin by width of antennal pedicel or more, usually much more, the distance between the fovea and eye much wider in the center than the ends of the fovea." Not obviously the case here.

Andrena prunorum prunorum (Andrena)

Washington, 12mm;
Complete hair bands;
T2 pitting dense;
T2 surface shiny;
Propodeal sculpturing 100%;
Corbicula incomplete;
Hind tibial spurs broadened at base.

= A. prunorum

Unambiguous. The spur with broadened base is shared with only one other Washington species (A. crataegi). Add complete T2 hair band and only A. prunorum remains.

Image from Bees of Canada, Packer Lab.

Other hallmarks of the species: "the usually red legs of both sexes and the coarsely rugatulopunctate mesepisterna of both sexes."

Andrena salicifloris (Trachandrena)

Triangle 100%, with enclosing carina;

Oregon, 12mm;
Clypeus shiny, impunctate midline;

Incomplete corbicula;
T2 pit spacing 1-3;
Pale scopal hairs;
Scutum with dense pitting;

Andrena fuscicauda
Andrena miranda
Andrena salicifloris

All matches are subgenus Trachandrena, characterized coarse sculpturing, an entirely rugose, enclosed propodeum, and fovea narrowed at the level of the antennal socket. DL does not unamnbiguously resolve species. So I consulted A Revision of the Bees of the Genus Andrena of the Western Hemisphere. Part VI. Subgenus Trachandrena (LaBerge). A. fuscicauda doesn't match this specimen in several respects. Differentiating miranda and salicifloris is harder.

miranda vs salicifloris: A. salcifloris and A. miranda split on a final couplet. A. salicifloris: mesoscutum with anterior third with punctures crowded, separated by mere ridges; vertex above lateral ocellus equals about one ocellar diameter.
A. miranda: Mesoscutum with anterior third with punctures discrete, separated by half to almost one puncture width; vertex above lateral ocellus usually distinctly longer than one ocellar diameter.
My specimen appears to have crowded punctures AND a long vertex. To quote LaBerge: "There is no doubt that misidentification of such specimens will continue and, indeed, it is often impossible to place some in the correct species with any confidence."

Above: Narrow lower fovea of Trachandrena.
Below: Scutum of another specimen.

Andrena vicina--Michigan (Melandrena)

Michigan, 13mm, May;
Clypeus shiny/dull;
Vertex ~1.8ocd;

Tibial hairs dark;
Abdominal hairs dark;
T2 hair band absent;
T2 puncate, 1-3 ;
Scutm punctate;
Trochanteral flocculus: variable;

Abundance of caution. Consider other MI Melandrena with dark hairs on tibia and T2-5 tergae:

A. vicina
A. nivalis

A. vicina
A. nivalis
A. regularis
A. carlini